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Sevastopol5Kazan17Q1668803920Odesa25embryology67Doctor of Natural Sciences106Tübingen555Pour le Mérite for Sciences and Arts order627Heidelberg664Order of Saint Stanislaus, 1st class708Order of St. Vladimir, 3rd class762Foreign Member of the Royal Society764Order of Saint Anna, 2nd class860biology926zoology972Order of Saint Anna, 1st class98919012,006